Oliver Smithies

Awards

1. 1990

   Canada Gairdner International Award

   Biology

   For the discovery, development and application of gel electrophoresis methods that allow the separation and identification of specific proteins and nucleic acids.

2. 1993

   Canada Gairdner International Award

   Biology

   For pioneering work in the use of homologous recombination to generate targeted mutations in the mouse.

3. 2001

   Lasker Award — Albert Lasker Basic Medical Research Award

   Biology

   For the development of a powerful technology for manipulating the mouse genome with exquisite precision, which allows the creation of animal models of human disease.

4. 2002/2003

   Wolf Prize — Medicine

   Biology

   for the development of procedures to manipulate mouse ova and embryos, which has enabled transgenesis and its applications in mice.

5. 2007

   Nobel Prize — Medicine

   Biology

   for their discoveries of principles for introducing specific gene modifications in mice by the use of embryonic stem cells